The entire imagery depicts a reversal of creation’s order. Humanity originally had been given the responsibility for exercising dominion over the whole of creation. But now the creation seizes power from the greatest of human empires and transforms it into a bestial wilderness. The animals will occupy the land, which is another way of pointing to a lack of human inhabitants.
14 Herds shall lie down in her midst, all kinds of beasts; even the owl and the hedgehog shall lodge in her capitals; a voice shall hoot in the window; devastation will be on the threshold; for her cedar work will be laid bare.